Present address:   Department of Biology, University of Bergen, P.O. Box 7800, 5020 Bergen, NORWAY

Tel:                         +47 – 55 58 44 35                               Fax:        +47 – 55 58 44 50

e-mail:                        Date of Birth:            May 16, 1956



B.Sc. with honours (Biological Oceanography, Fisheries) University of Michigan (USA)

Ph.D. University of Stirling (UK)


Positions held:

2003 – present: Professor, Department of Biology, University of Bergen


1988-2003: Research Fellow, Port Erin Marine Laboratory, University of Liverpool, Port Erin, IOM.

1985-Dec87: Assistant Professor (Lecturer), Environmental Sciences Center, Unity College, Unity, Maine, USA. Responsible for development of program and curriculum leading to a Bachelor's degree in Aquaculture. Teaching responsibilities included courses in vertebrate biology, zoology, physiology, ichthyology, fisheries science, aquaculture, and marine biology.

1982‑1985: Project Associate, Center for Limnology, University of Wisconsin, Madison, WI. Wisconsin Sea Grant project: Species Interactions and early life history dynamics of forage fish in Lake Michigan.

1976‑1979: Research Assistant, Mariculture Unit, Israel Oceanographic & Limnological Research Company, Eilat, ISRAEL. Nutrition, Diet preparation and testing, Cage culture design and construction, Broodstock management, Larval rearing, Food chain and hatchery techniques, Laboratory Assistant, Kinneret Limnological Laboratory, Israel Oceanographic and Limnological Research Company, Tiberias, ISRAEL. Zooplankton physiology and distributions, Primary and secondary production, Development of early feeding behaviour in Tilapia.


Research and Academic Experience

Assistant editor, Journal of Fish Biology (1993-1997)

Quality Assessor, Welsh Funding Council (Higher Education Funding Council for Wales) (1994-95)

Member, European Fish Ageing Network, European Union concerted action (1997-1999)

International Steering Committee, International Conference on Fish Otolith Research and Application

Co-convener. International Symposium on Ichthyoplankton Ecology. Annual Meeting of Fisheries Society of the British Isles 1997, guest editor of proceedings (Journal of Fish Biology, supplement 1997)

Conference Organising Committee, International Symposium on Flatfish Ecology (2000-2002), guest editor of proceedings, Journal of Sea Research 2003 & 2004

Review panel: Charles Lindberg Awards (1995-2000)

Expert review panel: INTAS (European Union) (2001-2008)

Review panel: Enterprise Ireland panel Basic Research Grant Programme (2001-2008)

Steering Committee: TACADAR (European Fish Ageing Network) Concerted Action 2003 – 2005

Editorial Board: ICES Journal of Marine Science (2005 – present)

Head of Aquaculture Biology Study Program, University of Bergen (2003 – present)

Committee leader: PhD examinations, appointments


External examiner (PhD)

University of Glasgow (1990), University of Aberdeen (1998), University of Plymouth (2000), University of Wales Aberystwyth (2004), University College Galway (2008), Galway-Mayo Institute of Technology (2008), University of Aberdeen (2008), University of East Anglia (2009), University College Dublin (2010), University of Southampton (2012)


Recent Grants awarded

EU Framework 5 “CODTRACE - Establishing traceability for cod (Gadus morhua): determining location of spawning and harvest” 2002 – 2004   

EU Framework 5 “IBACS - Integrated Approach to the Biological Basis of Age Estimation in Commercially Important Fish Species” 2002 – 2005 

EU Framework 5 “WESTHER - A multidisciplinary approach to the identification of herring (Clupea harengus L.) stock components west of the British Isles using biological tags and genetic markers” 2002-2005  

EU Accompanying Measures “FIFS Int’l Symposium on the Role of Flatfishes in Benthic Ecosystems” 2002–04 

EU - Framework 7 – “FishPopTrace” - The structure of fish populations and traceability of fish and fish products 2008 - 2011

EU - Framework 7 – “CalMarO” - Calcification of Marine Organisms - A Marie Curie ITN (Initial Training Network) concerned with understanding biomineralization and the effects of ocean acidification on calcifying organisms 2008 – 2012

NFR - HAVKYST  - PRIBASE: Pollution Risk and Impact Analysis for the Barents Sea Ecosystem 2009-2011

EEA - Managing effects of global climate on estuarine biodiversity and productivity, network project PT0010, Norway-the Netherlands-Portugal 2008-2012

NFR - The occurrence of skipped spawning and its importance for population dynamics in Northeast Arctic gadoids - 2009 – 2012


Professional Memberships:

American Fisheries Society: Early Life History Section, Fisheries Society of the British Isles, Marine Biological Association of the United Kingdom, Scottish Association for Marine Science, World Aquaculture Society


Recent Publications (since 2008) from 88 peer-reviewed


Rowlands WLl, Dickey-Collas M, Geffen AJ, Nash RDM. (2008) Diet overlap and prey selection through metamorphosis in Irish Sea cod (Gadus morhua), haddock (Melanogrammus aeglefinus) and whiting (Merlangius merlangus). Canadian Journal of Fisheries and Aquatic Sciences 65: 1297 – 1306.

Li L, Høie H, Geffen AJ, Heegaard E, Skadal J, and Folkvord A (2008)  HHBack-calculation of previous fish size based on tagged and marked Atlantic cod (Gadus morhua). Canadian Journal of Fisheries and Aquatic Sciences 65: 2496–2508 

Nash RDM, Kjesbu OS, Trippel EA, Finden H, and Geffen AJ. (2008) Potential variability in the paternal contribution to stock reproductive potential of Northeast Arctic cod (Gadus morhua). Journal of Northwest Atlantic Fishery Science 41: 71–83

Geffen AJ (2009) Advances in herring biology: from simple to complex, coping with plasticity and adaptability. ICES Journal of Marine Science 66: 1688–1695

van der Veer HW, Bolle LJ, Geffen AJ, Witte J IJ. (2009). Variability in transport of fish eggs and larvae. IV. Interannual variability in larval stage duration of immigrating plaice in the Dutch Wadden Sea. Marine Ecology Progress Series 390: 213–223  DOI: 10.3354/meps08176

Hufthammer AK, Høie H, Folkvord A, Geffen AJ, Andersson C, Ninnemann US (2010). Seasonality of human site occupation based on stable oxygen isotope ratios of cod otoliths. Journal of Archaeological Science, 37: 78–83DOI:10.1016/j.jas.2009.09.001

Watts PC, Kay SM, Wolfenden D, Fox CJ, Geffen AJ, Kemp SJ, and Nash RDM. (2010) Temporal patterns of fine-scale spatial genetic structure and effective population size in European plaice (Pleuronectes platessa L.) along the west coast of Scotland and the Irish Sea.  ICES Journal of Marine Science, 67: 607-616. DOI:10.1093/icesjms/fsp274

Cuveliers EL, Geffen AJ, Guelinckx J, Raeymaekers JAM, Skadal J, Volckaert FAM and Maes GE. (2010) Microchemical variation in juvenile sole (Solea solea) otoliths as a powerful tool for studying connectivity in the North Sea. Marine Ecology Progress Series 401: 211–220,

Therkildsen NO, Nielsen EE, Hüssy K, Meldrup D, Geffen AJ. (2010) Does DNA extraction affect the physical and chemical composition of historical cod (Gadus morhua) otoliths? ICES Journal of Marine Science 67: 1251-1259. DOI: 10.1093/icesjms/fsq016

Seljeset O, Vollset KW, Folkvord A, and Geffen AJ. (2010) The role of prey concentration and size range in the growth and survival of larval cod. Marine Biology Research 6: 251 – 262.

Higgins RM, Danilowicz BS, Balbuena JA, Daníelsdóttir AK, Geffen AJ, Meijer WG, Modin J, Montero F, Pampoulie C, Perdiguero D, Schreiber A, Stefánsson MO, Wilson B. (2010) Multi-disciplinary Fingerprints Reveal the Harvest Location of Cod (Gadus morhua) in the Northeast Atlantic. Marine Ecology Progress Series 404: 197–206 DOI: 10.3354/meps08492

Groison A-L, Suquet M, Cosson J, Mayer I, Severe A, Bouquet J-M, Geffen AJ, Utne-Palm AC, Kjesbu OS (2010) Sperm motility in European hake, Merluccius merluccius, and characterization of its spermatozoa concentration and volume, spermatocrit, osmolality and pH. Aquaculture 301: 31-36. DOI:10.1016/j.aquaculture.2010.01.026

Dickey-Collas M, Nash RDM, Brunel T, van Damme CJG, Marshall CT, Payne MR, Corten A, Geffen AJ, Peck MA, Hatfield EMC, Hintzen NT, Enberg K, Kell LT, Simmonds EJ (2010) Lessons learned from stock collapse and recovery of North Sea herring: a review. ICES Journal of Marine Science 67, 1875-1886. DOI:10.1093/icesjms/fsq033

Meier S, Morton CH, Nyhammer G, Grøsvik BE, Makhotin V, Geffen A, Boitsov S, Kvestad KA, Bohne-Kjersem A, Goksøyr A, Folkvord A, Klungsøyr J, Svardal A (2010) Development of Atlantic cod (Gadus morhua) exposed to produced water during early life stages: Effects on embryos, larvae, and juvenile fish. Marine Environmental Research 70: 383-394. DOI:10.1016/j.marenvres.2010.08.002

Ciotti, B. J., Targett, T. E., Nash, R. D. M., Batty, R. S., Burrows, M. T. and Geffen, A. J. (2010) Development, validation and field application of an RNA-based growth index in juvenile plaice Pleuronectes platessa. Journal of Fish Biology, 77: 2181–2209. DOI:10.1111/j.1095-8649.2010.02786.x.

Geffen AJ, Nash RDM, Dau K, Harwood AJP (2011) Sub-cohort dynamics of O-group plaice, Pleuronectes platessa L., in the Northern Irish Sea: settlement, growth and mortality.  Journal of Experimental Marine Biology and Ecology. 400 (1-2):108-119.  DOI:10.1016/j.jembe.2011.02.030

Geffen AJ, Høie H, Folkvord A, Hufthammer, AK, Andersson, C, Ninemann U, Pedersen, RB, Nedreaas K (2011) High latitude climate variability and its effect on fishery resources as revealed by fossil otoliths. ICES Journal of Marine Science. 68: 1081-1089.   DOI:10.1093/icesjms/FSR017

Kennedy J, Skjæraasen JE, Nash RDM, Slotte A, Geffen AJ, Kjesbu OS (2011) Evaluation of the frequency of skipped spawning in Norwegian spring-spawning herring, Journal of  Sea Research 65: 327-332.  DOI:10.1016/j.seares.2011.01.003

Geffen AJ, Nash RDM, Dickey-Collas M (2011) Characterisation of herring populations to the west of the British Isles: an investigation of mixing between populations based on otolith microchemistry. ICES Journal of Marine Science. 68: 1447-1458.  DOI: 10.1093/icesjms/FSR051

Meier S, Morton HC, Andersson E, Geffen AJ, Taranger GL, Larsen M, Petersen M, Djurhuus R, Klungsøyr J, Svardal A. (2011) Low-dose exposure to alkylphenols adversely effects the sexual development of Atlantic cod (Gadus morhua): Acceleration of the onset of puberty and delayed seasonal gonad development in mature female cod. Aquatic Toxicology 105: 136-150.  DOI: 10.1016/j.aquatox.2011.06.003

Geffen AJ, Nash RDM (2012) Egg development rates for use in Egg Production Method (EPMs) and beyond. Fisheries Research  117–118: 48–62    DOI:10.1016/j.fishres.2011.01.002

Nash RDM, Geffen AJ (2012) Mortality in the early life-history of fish: What can we learn from European plaice (Pleuronectes platessa L.)? Journal of Marine Systems. 93: 58–68

Frommel AY, Maneja R, Lowe D, Malzahn AM, Geffen AJ, Folkvord A, Piatkowski U, Reusch TBH, Clemmesen C. (2012) Severe tissue damage in Atlantic cod larvae under increasing ocean acidification. Nature Climate Change 2: 42–46.   DOI:10.1038/nclimate1324

Morales-Nin B, Geffen AJ, Pérez-Mayol S, Palmer M, González-Quirós R, Grau A. (2012) Seasonal and ontogenic migrations of meagre (Argyrosomus regius) determined by otolith geochemical signatures. Fisheries Research 127– 128: 154– 165.   DOI:10.1016/j.fishres.2012.02.012

Geffen AJ. (2012) Otolith oxygen and carbon stable isotopes in wild and laboratory-reared plaice (Pleuronectes platessa). Environmental Biology of Fishes DOI: 10.1007/s10641-012-0033-2

Skjaeraasen JE,  Nash RDM, Korsbrekke K, Fonn M, Nilsen T, Kennedy J, Nedreeas KH, Thorsen A, Witthames PR, Geffen AJ, Høie H, Kjesbu OS. (2012) Frequent skipped spawning in the world's largest cod population. Proceedings of the National Academy of Sciences. DOI:10.1073/pnas.1200223109

Chang M-Y, Geffen AJ. (2012) Taxonomic and environmental influences on fish otolith microchemistry.  Reviews in Fish and Fisheries DOI: 10.1111/j.1467-2979.2012.00482.x

Chang M-Y, Geffen AJ, Kosler J, Dundas SH, FishPopTrace Consortium,  Maes GE (2012)  The effect of ablation pattern on LA-ICPMS analysis of otolith element composition in hake, Merluccius merluccius. Environmental Biology of Fishes DOI: 10.1007/s10641-012-0065-7

Exadactylos A, Malandrakis EE, Panagiotaki P, Geffen AJ (in press) The development of size variation in Dover sole, Solea solea and turbot, Scophthalmus maximus: genetic variability between different geographic and among year class farmed strains. Aquaculture Research DOI: 10.1111/j.1365-2109.2012.03196.x

Wang C-H, Geffen AJ, Nash RDM. (in press) Geographical variation in the chemical composition of veined squid Loligo forbesi statoliths. Zoological Studies. (Accepted 13.3.2012)

Maneja RH, Frommel AY, Browman HI, Clemmesen C, Geffen AJ, Folkvord A, Piatkowski U,  Durif CMF, Bjelland R, Skiftesvik AB (in press) The swimming kinematics of larval Atlantic cod, Gadus morhua L., are resilient to elevated seawater pCO2.  Marine Biology

Dolphin AE, Dundas SH, Košler J, Tvinnereim HM, Geffen AJ (in press) A comparison of techniques for measuring the trace element content of human teeth: laser ablation ICP-MS versus solution ICP-MS of micromilled enamel. International Journal of Analytical and Bioanalytical Chemistry ISSN 2231-5012  (Accepted September 2012)

Morey G, Morales-Nin B, Riera F, Grau A, Geffen AJ, Pérez-Mayol S, Chang M-Y, Grau AM. (in press) Mediterranean cod (Gadus morhua): a surprising emigrant submitted to Marine Ecology Progress Series doi: 10.3354/meps09936